Physical–biological coupling in spore dispersal of kelp forest macroalgae

The physical–biological linkages controlling the dispersal of spores produced by macroalgae that reside in kelp forests are complicated and laced with feedbacks. Here we discuss the fundamental elements of these interactions. Biological considerations include spore swimming and sinking speeds, their periods of viability in the plankton, and the height of spore release above the seafloor, which together determine the durations over which spores can be swept by horizontal currents before they contact the seafloor. Morphologies and material properties of canopy forming kelps may also influence the drag exerted on passing waters by the kelps, the plants' ability to persist in the face of rapid flows, and thereby the degree to which impinging currents are redirected around, or slowed within, kelp forests. Macroalgal life histories, and the size of spore sources as controlled by the dimensions of kelp forests and the density and fecundity of individuals within them, influence effective dispersal distances as well. Physical considerations encompass the mean speed, direction, and timescales of variability of currents relative to spore suspension times, the interaction of surface gravity waves with currents in producing turbulence in the benthic boundary layer, wind-driven surface mixing, water stratification, and shoreline bathymetry and substratum roughness, all of which can affect the interplay of vertical and horizontal transport of macroalgal spores. Intricate within-forest processes may induce attenuation of current speeds and consequent reductions in seabed shear, along with simultaneous production of small-scale turbulence in kelp wakes. Slower mean currents and smaller eddy scales in turn may attenuate vertical mixing within forests, thus extending spore suspension times. Further complexities likely arise due to changes in the relative rates of horizontal and vertical dispersion, modifications to the overall profiles of vertical mixing, and the creation of fine-scale secondary flows around kelp individuals and substratum features. Under conditions of more rapid currents, submergence of the surface canopy and the establishment of skimming flows at the canopy–fluid interface may introduce additional coherent flow structures that alter rates of fluid exchange to and from the forest. Many of these coupled physical–biological processes are just beginning to be examined in a rigorous fashion in kelp forests, but their potential importance is clear.     

Modification of the Highway Capacity Manual two‐lane highway analysis procedure for Spanish conditions

The US Highway Capacity Manual (HCM) methodology is used in Spain to evaluate traffic operation and quality of service. The effect of passing manoeuvre on two‐lane highway operational performance is considered through adjustment factors to average travel speeds and percent time spent following. The procedure is largely based on simulations in TWOPAS and passing behaviours observed during US calibrations in the 1970s. It is not clear whether US driving behaviour and vehicles' performance are comparable with Spanish conditions. The objective of this research is to adapt the HCM 2010 methodology to Spanish driver behaviour, for base conditions (i.e. no passing restrictions). To do so, TWOPAS was calibrated and validated based on current Spanish passing field data. The calibration used a genetic algorithm. The case study included an ideal two‐lane highway with varying directional traffic flow rate, directional split and percentage of trucks. The updated methodology for base conditions is simpler than the current HCM 2010 and does not rely on interpolation from tables. Copyright © 2016 John Wiley & Sons, Ltd.