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1.
An upgraded and revised physically–biologically coupled, nested 3D model with 4 km grid size is applied to investigate the seasonal carbon flux and its interannual variability. The model is validated using field data from the years for which the carbon flux was modelled, focussing on its precision in space and time, the adequacy of the validation data, suspended biomass and vertical export. The model appears to reproduce the space and time (± 1 week and 10 nautical miles) distribution of suspended biomass well, but it underestimates vertical export of carbon at depth. The modelled primary production ranges from 79 to 118 g C m 2 year 1 (average 93 g C m 2 year 1) between 4 different years with higher variability in the ice-covered Arctic (± 26%) than in the Atlantic (± 7%) section. Meteorological forcing has a strong impact on the vertical stratification of the regions dominated by Atlantic water and this results in significant differences in seasonal variability in primary production. The spatially integrated primary production in the Barents Sea is 42–49% greater during warm years than the production during the coolest and most ice-covered year.  相似文献   

2.
Oil sac volume, gonad size and moulting patterns were investigated in the copepod Calanus euxinus inhabiting deep and shallow zones of the Black Sea and penetrating into the Marmara Sea. In summer the C. euxinus population in deep layers of the Black Sea was dominated by pre-diapause and diapausing postmoult copepodite stage V (CV) with small sexually undifferentiated gonads and mean lipid content of 14.1 ± 6.0% of body volume. The lipid content of deep-living females was 7.2 ± 4.2% of body volume. At the same time, intermoult and premoult CV with enlarged gonads and low lipid content (7.7 ± 5.1% of body volume) and females with oil sac volume of 1.4 ± 1.0% were found at shallow stations. Premoult CV with oil volume of 0.6 ± 0.8% and mature females with little visual evidence of substantial lipid storage dominated in the Marmara Sea. The differences in moulting patterns and oil sac volumes of C. euxinus from deep zones and shallow regions suggest that vertical migrations to the oxygen minimum zone (OMZ) are necessary for formation of large lipid reserves providing high reproductive potential of this species. On the basis of an energy balance model it was shown that under low phytoplankton concentration of about 30 μg C l 1 preadults and adults migrating to the OMZ could accumulate lipids (up to 5% of body energy content daily), in contrast to copepods constrained to shallow oxic water columns of the Black Sea and from the Marmara Sea.  相似文献   

3.
The large quantities of particles delivered by the Mackenzie River to the coastal Beaufort Sea (Arctic Ocean) have implications for the spatial distribution, composition and productivity of its bacterial communities. Our objectives in this study were: (1) to assess the contribution of particle-associated bacteria (fraction  3 µm) to total bacterial production and their relationships with changing environmental conditions along a surface water transect; (2) to examine how particle-based heterotrophy changes over the annual cycle (Nov 2003–Aug 2004); and (3) to determine whether particle-associated bacterial assemblages differ in composition from the free-living communities (fraction < 3 µm). Our transect results showed that particle-associated bacteria contributed a variable percentage of leucine-based (BP-Leu) and thymidine-based (BP-TdR) bacterial production, with values up to 98% at the inshore, low salinity stations. The relative contribution of particle-associated bacteria to total BP-Leu was positively correlated with temperature and particulate organic material (POM) concentration. The annual dataset showed low activities of particle-associated bacteria during late fall and most of the winter, and a period of high particle-associated activity in spring and summer, likely related to the seasonal inputs of riverine POM. Results from catalyzed reporter deposition for fluorescence in situ hybridization (CARD-FISH) confirmed the dominance of Bacteria and presence of Archaea (43–84% and 0.2–5.5% of DAPI counts, respectively), which were evenly distributed throughout the Mackenzie Shelf, and not significantly related to environmental variables. Denaturing gradient gel electrophoresis (DGGE) revealed changes in the bacterial community structure among riverine, estuarine and marine stations, with separation according to temperature and salinity. There was evidence of differences between the particle-associated and free-living bacterial assemblages at the estuarine stations with highest POM content. Particle-associated bacteria are an important functional component of this Arctic ecosystem. Under a warmer climate, they are likely to play an increasing role in coastal biogeochemistry and carbon fluxes as a result of permafrost melting and increased particle transport from the tundra to coastal waters.  相似文献   

4.
We present a coupled sea ice–ocean-biological (including ice algae) model in the Arctic Ocean. The 1D model was developed and implemented on the Canadian Beaufort Sea shelf to examine the importance of different physical processes in controlling the timing and magnitude of primary production and biogenic particle export over an annual cycle (1987). Our results show that the snow and sea ice cover melt and/or break-up controls the timing of the phytoplankton bloom but primary producers (ice algae and phytoplankton) on the outer shelf are essentially nutrient limited. The total annual primary production (22.7 to 27.7 g-C m? 2) is thus controlled by nutrient “pre-conditioning” in the previous fall and winter and by the depth of wind mixing that is controlled in part by the supply of fresh water at the end of spring (ice melt or runoff). The spring bloom represents about 40% of the total annual primary production and occurs in a period of the year when sampling is often lacking. Time interpolation of observed values to obtain total annual primary production, as done in many studies, was shown to lead to an underestimation of the actual production. Our simulated ratios of export to primary production vary between 0.42 and 0.44.  相似文献   

5.
Surfical sediments within Corunna Lake, a moderate size Intermittently Closed and Open Lake Lagoon (ICOLL), were examined for solid phase nutrient concentrations (TN, TP, TOC,) and solute exchange rates between the sediment and water column (O2, NO3–N, NH4–N, FRP, and N2). The surfical sediments in Corunna Lake contained high concentrations of TN (5 mg/g dry mass), total phosphorus (0.6 mg/g dry mass), and TOC (~ 5% dry mass). The carbon stable isotope ratio (δ13C) and TOC:TN ratios (δ13C ~ ? 24, TOC:TN ~ 11–14) demonstrated that the composition of the organic matter in the sediment was a mixture derived primarily of degraded planktonic matter. The close association between TP and Fe concentrations highlighted the potential role Fe plays in mediating Filterable Reactive Phosphorus (FRP) concentrations in the water column of Corunna Lake. In situ benthic chamber incubations were used to measure benthic fluxes. Solute exchange rates between the sediment and water column in Corunna Lake were similar to other reported studies (O2 = ? 469 to ? 1765 µmol m? 2 h? 1, NH4–N = 0.1–63 µmol m? 2 h? 1, NO2/NO3–N = 0 µmol m? 2 h? 1, FRP = ? 4–1.6 µmol m? 2 h? 1and N2 = 12–356 µmol m? 2 h? 1). As more carbon was deposited and mineralized the efficiency of the bacterial population to denitrify nitrogen in the sediment decreases. The linkage between land use and benthic biogeochemistry was also explored. A dairy farm exists in the middle catchment of Corunna Lake, and the receiving bay sediment consistently demonstrated the highest oxygen consumption rates in winter and spring (? 1408 µmol m? 2 h? 1 in winter, ? 1691 µmol m? 2 h? 1 in spring) and lowest denitrification efficiencies during summer (~ 3%). Nitrate/nitrite fluxes were not observed during any of the chamber incubations, with the concentrations of nitrate/nitrite being below detection limits (< 10 μg/L). Seasonal changes influenced the rates of solute exchange between the sediment and water column. Critical measures of solute exchange for NH4–N and biogenic N2 indicated that seasonal temperature changes play a significant role in mediating the reaction rates of sedimentary based biogeochemical processes. Measurable FRP fluxes were small but greater in the benthic sediments which received higher carbon inputs. Sediments have a high capacity to adsorb P which is released as sediment oxygen demand increases as a result of increases in labile carbon loads.  相似文献   

6.
Surface seawater samples were taken in the framework of the GEOTRACES program on “POLARSTERN” expedition ANT XXIII/1 in the Eastern Atlantic in 2005 to study the distribution of the trace elements Hg (mercury), Pb (lead), Cd (cadmium), Cu (copper), Ni (nickel), Zn (zinc), Co (cobalt), Mn (manganese), Fe (iron), and Al (aluminium). With the exception of Hg, results were compared to earlier datasets from 1989 to 1990. The particulate fraction averaged over the transect was calculated to be 49% for Cd, 23% for Mn and 50% for Fe indicating a release of these TEI's (trace elements and their isotopes) from a leachable SPM fraction in the stored and acidified samples.Total Pb concentrations ranged between 5 and 20 pmol kg? 1 in 2005 with highest values in the ITCZ (intertropical convergence zone). In 1989 Pb concentrations were twice as high in the region of the ITCZ, while by a factor of 10–15 higher values were obtained in the North Atlantic.Total Cd and Co are dominated, by different seasonal upwelling regimes (Equatorial upwelling, Guinea Dome, Angola Dome).Total Cu, Ni, Fe, Mn and Al show nearly identical concentrations in 1990 and 2005. For total manganese and aluminium strong maxima (3–4 nmol kg? 1 and 55 nmol kg? 1 respectively) are observed between 23°N and 0°, while the Fe maximum (6–9 nmol kg? 1) is located at 7°N. Total Hg concentrations ranged between 0.5 and 4.5 pmol kg? 1.  相似文献   

7.
An empirical algorithm has been developed to compute the sea surface CO2 fugacity (fCO2sw) in the Bay of Biscay from remotely sensed sea surface temperature (SSTRS) and chlorophyll a (chl aRS) retrieved from AVHRR and SeaWiFS sensors, respectively. Underway fCO2sw measurements recorded during 2003 were correlated with SSTRS and chl aRS data yielding a regression error of 0.1 ± 7.5 µatm (mean ± standard deviation). The spatial and temporal variability of air–sea fCO2 gradient (ΔfCO2) and air–sea CO2 flux (FCO2) was analyzed using remotely sensed images from September 1997 to December 2004. An average FCO2 of ? 1.9 ± 0.1 mol m? 2 yr? 1 characterized the Bay of Biscay as a CO2 sink that is suffering a significant long-term decrease of 0.08 ± 0.05 mol m? 2 yr? 2 in its capacity to store atmospheric CO2. The main parameter controlling the long-term variability of the CO2 uptake from the atmosphere was the air–sea CO2 transfer velocity (57%), followed by the SSTRS (10%) and the chl aRS (2%).  相似文献   

8.
High Reynolds number flows (Re = 1 × 106, 2 × 106 and 3.6 × 106, based on the free stream velocity and cylinder diameter) covering the supercritical to upper-transition flow regimes around a two-dimensional (2D) smooth circular cylinder, have been investigated numerically using 2D Unsteady Reynolds-Averaged Navier–Stokes (URANS) equations with a standard high Reynolds number k ? ? turbulence model. The objective of the present study is to evaluate whether the model is applicable for engineering design within these flow regimes. The results are compared with published experimental data and numerical results. Although the k ? ? model is known to yield less accurate predictions of flows with strong anisotropic turbulence, satisfactory results for engineering design purposes are obtained for high Reynolds number flows around a smooth circular cylinder in the supercritical and upper-transition flow regimes, i.e. Re > 106. This is based on the comparison with published experimental data and numerical results.  相似文献   

9.
《Journal of Marine Systems》2006,59(1-2):143-158
Effects of stocking density on seston dynamics and filtering and biodeposition by the suspension-cultured Zhikong scallop Chlamys farreri Jones et Preston in a eutrophic bay (Sishili Bay, northern China), were determined in a 3-month semi-field experiment with continuous flow-through seawater from the bay. Results showed that the presence of the scallops could strongly decrease seston and chlorophyll a concentrations in the water column. Moreover, in a limited water column, increasing scallop density could cause seston depletion due to scallop's filtering and biodeposition process, and impair scallop growth. Both filtration rate and biodeposition rate of C. farreri showed significant negative correlation with their density and positive relationship with seston concentration. Calculation predicts that the daily removal of suspended matter from water column by the scallops in Sishili Bay ecosystem can be as high as 45% of the total suspended matter; and the daily production of biodeposits by the scallops in early summer in farming zone may amount to 7.78 g m 2, with daily C, N and P biodeposition rates of 3.06 × 10 1, 3.86 × 10 2 and 9.80 × 10 3 g m 2, respectively. The filtering and biodeposition by suspension-cultured scallops could substantially enhance the deposition of total suspended particulate material, suppress accumulation of particulate organic matter in water column, and increase the flux of C, N and P to benthos, strongly enhancing pelagic–benthic coupling. It was suggested that the filtering-biodeposition process by intensively suspension-cultured bivalve filter-feeders could exert strong top-down control on phytoplankton biomass and other suspended particulate material in coastal ecosystems. This study also indicated that commercially suspension-cultured bivalves may simultaneously and potentially aid in mitigating eutrophication pressures on coastal ecosystems subject to anthropogenic N and P loadings, serving as a eutrophic-environment bioremediator. The ecological services (e.g. filtering capacity, top-down control, and benthic–pelagic coupling) functioned by extractive bivalve aquaculture should be emphasized in coastal ecosystems.  相似文献   

10.
This paper provides an in-depth study of residual stress distributions found in stiffened steel plate structures, such as those typically used in ship hulls. The effect of stiffener spacing on the distribution of residual stress components was studied. The welding heat input was also varied between high and moderate to study the effect of heat input level on residual stress distributions. Four specimens, resembling typical stiffened steel plate structures used in ship hulls were built and tested. Steel plates of 9.5 mm thickness were stiffened by welding L127 × 76 × 9.5 steel angles. The test was completed using the neutron diffraction method. The three normal components of residual stress were obtained in this study. It was found that a lower heat input results in higher tensile residual stress and that there exists a critical stiffener spacing somewhere beyond 250 mm that creates a maximum tensile residual stress value near the welded connection.  相似文献   

11.
This work estimates new regionalized empirical parameterizations for preformed alkalinity (ATo) and the CO2 air–sea disequilibrium (?Cdis). Both are key terms for the computation of anthropogenic CO2 in the back-calculation methods. Data from the subsurface layer (75–180 m depth range) covering an area from North to South and from 19°E to 67.5°W (Pacific and Indian oceans) were taken from GLODAP (The Global Ocean Data Analysis Project) database. The subsurface layer is proved as a reliable reference for representing the main characteristics of the different water masses of the oceans. Besides, handing data from the two ocean basins altogether makes the new parameterizations of ATo and ?Cdis to be more globally consistent. Nevertheless, each ocean basin, at least in some regions, has different oceanographic characteristics based on its proper dynamical processes and water masses formation. In order to maintain each ocean basin ‘identity’ the whole domain was divided in six different regions (two of them sharing waters from Pacific and Indian oceans) and parameterizations in each region for both terms were obtained. Previously, data were transformed into a grid of 4°lat. × 5°lon. and the results obtained from the parameterizations were visualized and compare with pCO2 climatologies. From the comparisons with previous ?Cdis estimations good results are obtained showing the reliability and robustness of the new regionalized empiric parameterizations.  相似文献   

12.
《Journal of Marine Systems》2006,59(1-2):111-119
The seasonal and spatial variations of chlorophyll concentrations, Sea Surface Temperature (SST), wind fields and wind-induced Ekman pumping in the Sulu Sea are investigated using a set of new remote sensing measurements from October 1997 to December 2004. The results show the seasonality of chlorophyll, wind fields and SST and reveal the phytoplankton blooming events in the Sulu Sea basin during the northeast monsoon season. In summer, chlorophyll concentrations were relatively low (< 0.2 mg/m3) and distributed uniformly throughout the basin with a narrow belt of high chlorophyll concentrations along the coastal waters, particularly the coasts of Borneo and of the Sulu Archipelago. In winter, chlorophyll concentrations increased (> 0.2 mg/m3) throughout the entire basin, and phytoplankton bloomed southward to the central basin, while chlorophyll concentrations reached high levels (1 mg/m3) in the center of the blooms. One peak was observed during the northeast monsoon season each year. SSTs have significant negative correlations with chlorophyll concentrations; i.e., high and uniformly distributed in summer but lower with an obvious tongue of cold waters southward to the central basin in winter. The seasonal variation of chlorophyll concentrations and SST distribution were associated with the seasonally reversing monsoon. The winter phytoplankton blooming and the tongue of the cold waters were correlated to the vertical upwelling cold and nutrient-rich waters drawn by the northeast wind, with the center of the blooms and the location of cold tongues coinciding with the maximum of the wind speeds and the Ekman pumping velocities.  相似文献   

13.
14.
Downward fluxes of organic biodeposits under suspended mussel culture cause benthic impacts such as microbial mat production. Quantifying sediment erosion in these coastal ecosystems is important for understanding how fluxes of organic matter and particulates contribute to benthic–pelagic coupling. Critical shear velocity (u?), erosion rates and particle size distributions of resuspended sediment were measured at two sites; an impacted muddy site with extensive mussel culture (site 1), and a coarser sandier site with less mussel influence (site 2), using a new method for assessing sediment erosion at Tracadie Bay, Prince Edward Island in August 2003. Shear forces were generated by vertically oscillating a perforated disc at controlled frequencies. These forces correspond to shear velocity, using a re-designed and calibrated Particle Erosion Simulator. Undisturbed sediment cores obtained by divers and grab (sub-cored using a Plexiglas? cores) were exposed to shear stress to compare differences between collection methods. Microbial mats were present at site 1 which initially biostabilized sediment against erosion due to ‘armoring’ of the sediment, but onset of erosion was abrupt once these mats failed. Erosion sequences at site 2 (without mat cover) were smoother resulting in less material being eroded. Mean mass of material eroded was 47 and 23 g m? 2 min? 1 at sites 1 and 2 respectively. Mat area cover and shear velocity was strongly related. Critical shear velocities varied between 1.70 and 1.77 cm s? 1, with no obvious differences between location or collection method, so sediments from these two contrasting sites had identical mean critical shear velocities. Significant differences existed in the concentrations of chlorophyll a, colloidal and bulk carbohydrates, between mats and bare sediment from site 1. Particle sizes measured by videography of resuspended sediment at different shear velocities ranged from 100 μm (the minimum diameter capable of being detected by the system), to large mat fragments of 1700 μm for both sites. These results provide evidence of the relevance of using a portable erosion device to indicate how sediment erodability is affected by mussel–microbial relationships.  相似文献   

15.
The spatial distribution of chlorophylls and carotenoids was recorded throughout the Gulf of Gabes (South Ionian Sea) in March 2007, and was related to patterns of the physical structure and the nutrient concentrations.Two distinct water masses were identified based on the temperature and salinity (TS) analysis: a cool and less salty Modified Atlantic Water (MAW) and a saltier Mediterranean Mixed Water (MMW). There was no significant difference in the mean nitrogen and phosphate concentrations between MMW and MAW, although the silica values were significantly higher in MAW. The Integrated chlorophyll a mean value was about 4 mg m? 2, with a maximum of 13 mg m? 2 at MAW stations.Higher Chlorophyll a records in typical MAW stations were mainly due to chlorophytes, which contributed up to 58% of the pigments concentrations in the MAW and about 46% in the MMW. The contribution of chlorophytes to total Chlorophyll a was found to be relatively stable throughout the water column. The contribution of diatoms, which were twofold higher in the MMW than in the MAW, did not exceed 17% of chlorophyll a and was mainly due to subsurface maxima. The chlorophytes, pelagophytes, prymnesiophytes and cryptophytes all together accounted for more than 77% of total chlorophyll a in the MAW and about 67% in the MMW.There were statistically significant differences between MMW and MAW in the pigment contribution of cyanobacteria and pelagophytes. These two taxa accounted for 13% and 24% of chlorophyll a respectively in the MAW and MMW indicating that these differences concerned phytoplankton classes at relatively low contributions to total chlorophyll a.  相似文献   

16.
Experimental work carried out at 1:60 scale in a wave flume assessed the pitch motion and anchor loading of 3 articulated tower installations in 50 m water depth while being exposed to north Atlantic storms with Hs of 15.2 m and Tp of 18.4 s. The three installations differ only in that their mass and buoyancy characteristics provide a natural period in pitch at equilibrium of 13 s, 20 s and 34 s respectively. It is verified that the dominant behaviour can be simulated by a relatively simple mathematical model, allowing the critical parameters of peak anchor loads and pitch angles to be calculated and extrapolated to full scale. It is demonstrated from the experimental and simulation results that the mass characteristics of a non surface piercing tower can be used to offset some of the challenges of moving to shallow water. If done correctly, it is possible to keep horizontal anchor loads under control and reduce vortex-induced transverse loading at the expense of increased pitch motions. Overall, the use of articulated tower installations in water depths of 50 m would appear to be technically feasible, even in exposed areas. The limitations on the size of such structures and the consequences of the resulting pitch accelerations and induced anchor loads are the subject of further study. It is proposed that the model verified herein can be used to further assess their potential at delivering viable wave power position mooring systems.  相似文献   

17.
In this study, a model for predicting chloride penetration in fly ash concrete under long-term exposure in a marine environment is developed. The empirical model was based on 2-, 3-, 4-, and 5-year investigation of concretes in a marine site. Regression analysis of the data was carried out by applying Fick's second law of diffusion to generate an empirical formula for predicting chloride concentration in concrete. The model uses the water to binder (W/B) ratio, fly ash content, distance from the concrete surface, and exposure time. Model validation revealed that the predicted chloride concentration levels were within a ±25% error margin (R2 = 0.91 ? 0.99) in the samples used to develop the model. The model was also verified using data from previous laboratory and field studies. Most predicted chloride concentration levels were within a ±30% margin of error from field samples. The model also predicted the strong effect of fly ash and W/B ratio on reducing chloride diffusion in concrete. Results clearly indicated that a high volume fly ash replacement (up to 50% by weight of binder) and a low W/B ratio will yield good chloride resistance in concrete under long-term exposure in a marine environment.  相似文献   

18.
Depth profiles of heterotrophic bacteria abundance were measured weekly over a 6-month period from December to May in Franklin Bay, a 230 m-deep coastal Arctic Ocean site of the southeastern Beaufort Sea. Total bacteria, low nucleic acid (LNA) and high nucleic acid (HNA) bacteria abundances were measured using flow cytometry after SYBR Green I staining. The HNA bacteria abundance in surface waters started to increase 5–6 weeks after phytoplankton growth resumed in spring, increasing from 1 × 105 to 3 × 105 cells mL− 1 over an 8-week period, with a net growth rate of 0.018 d− 1. LNA bacteria response was delayed by more than two months relative to the beginning of the phytoplankton biomass accumulation and had a lower net growth rate of 0.013 d− 1. The marked increase in bacterial abundance occurred before any significant increase in organic matter input from river discharge (as indicated by the unchanged surface water salinity and DOC concentrations), and in the absence of water temperature increase. The abundance of bacteria below the halocline was relatively high until January (up to 5 × 105 cells mL− 1) but then decreased to values close to 2 × 105 cells mL− 1. The three-fold bacterial abundance increase observed in surface waters in spring was mostly due to HNA bacteria, supporting the idea that these cells are the most active.  相似文献   

19.
《Marine Structures》2004,17(1):1-27
This paper presents a combined experimental–numerical procedure for development and calibration of macroscopic crack propagation criteria in large-scale shell structures. A novel experimental set-up is described in which a mode-I crack can be driven 400 mm through a 20(+) mm thick plate under fully plastic and controlled conditions. The test specimen can be deformed either in combined in-plane bending and extension or in pure extension. Experimental results are described for 5 and 10 mm thick aluminium and steel plates. By performing an inverse finite-element analysis of the experimental results where the simulated crack growth is forced to correspond to the experimental observations, empirical criteria for ductile crack propagation emerge very clearly. Using the experiments with edge crack specimens (ECS) in combined in-plane bending and extension, crack propagation criteria are developed for steel and aluminium plates, mainly as curves showing the critical element deformation versus the shell element size. These derived crack propagation criteria are then validated against a separate set of experiments considering centre crack specimens (CCS) which have a different crack-tip constraint. The applicability of the often-used equivalent strain criterion is discussed versus a more rationally based criterion which takes into account the stress tri-axiality. A large-scale grounding experiment is also simulated showing very good agreement with measurements. The performance of the proposed model is in general good and it is believed that the presented results and experimental–numerical calibration procedure can be of use in practical finite-element simulations of collision and grounding events with the use of shell elements. As discussed, the paper provides a clean framework for further development of macroscopic crack propagation criteria in large-scale plate structures.  相似文献   

20.
The distribution and diet of larval and juvenile Arctic cod (Boreogadus saida) were studied during summer 2005 in the coastal Canadian Beaufort Sea. A total of 275 individuals were captured and the highest abundance was observed at station depths of 20–30 m. This corresponds well with the location of the frontal zone where the Mackenzie River plume water and open sea water meet. Diet examinations were performed on 220 Arctic cod, which were found undamaged from sampling. We observed a gradual decrease in prey number per fish and increase in prey size as larvae grew which corresponded to a shift from Rotifera and nauplii towards larger copepodid stages. However, at all sizes, the larvae remain generalists and feed on a broad range of organisms. Environmental changes due to climate warming could have a two-fold impact on fish larvae feeding in the studied region. First, the potential for increased primary production may lead to increased zooplankton production that may impact the feeding and nutrition positively. On the other hand, greater discharge of turbid water from the Mackenzie River may reduce light penetration in the water column that may negatively influence the ability of visual predators to successively forage.  相似文献   

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