Few marine phytoplankton have heteromorphic life cycles and also often dominate the ecosystems in which they occur. The class Prymnesiophyceae contains a notable exception: the genus Phaeocystis includes three species that form gelatinous colonies but also occur within their ranges as solitary cells. Phaeocystis antarctica and P. pouchetii are exclusively high latitude taxa, and are notable for regionally tremendous blooms of the colony stage. P. globosa occurs circumglobally, yet its colony blooms primarily are confined to colder waters within its range. Three additional species are warm water forms that have been reported only as solitary cells or loose aggregations that bear little resemblance to the organized colonies of the other taxa. Interpretation of existing data indicates that resource availability (light, temperature and nutrients) by itself is not sufficient to explain this distinction between cold-water colony-forming taxa and warm water solitary cell taxa, nor why colony development in P. globosa is essentially a spatially restricted phenomenon within a much broader geographic range. Colony development by P. globosa in situ has been observed at temperatures ≥20 °C, but only rarely and generally under conditions of seasonally or anthropogenically elevated nutrient supply. Data presented here demonstrate colony development at 20–22 °C in natural plankton communities from oligotrophic waters that were pre-screened through 63 μm mesh (i.e. lacking mesozooplankton and large microzooplankton), but not in unscreened communities containing microzooplankton and >63 μm zooplankton. Reduction of colony proliferation at higher temperatures by mesozooplankton grazing remains as an intriguing possibility that is consistent with available evidence to help explain differences in latitudinal extent of in situ colony development. These data are interpreted within a theoretical framework regarding the potential advantages and disadvantages of the two life cycle stages. 相似文献
To identify the potential relationship between Pheaocystis globosa bloom conditions and seawater properties, a hydrobiological survey was performed in the inshore waters of the Eastern English Channel over the course of the phytoplankton spring bloom. Chlorophyll concentration, auto- and hetero/mixotrophic composition of protists and standing stock, and seawater viscosity were measured weekly from March to June 2004. The decline of the bloom is characterized by a massive foam formation in the turbulent surf zone. Before foam formation, seawater viscosity significantly increased, showing a significant positive correlation with chlorophyll concentration. In contrast, after foam formation this correlation was negative, seawater viscosity kept increasing despite a sharp decrease in chlorophyll concentrations. No significant correlation has been found between seawater viscosity and the composition of the phytoplankton assemblages observed during the survey. However, significant positive correlations have been found between seawater viscosity and both the size and the abundance of P. globosa colonies. From the correlation patterns observed between chlorophyll concentration and seawater viscosity, we suggest that the rheological properties of seawater are mainly driven by extracellular materials associated with colony formation and maintenance rather than by cell composition and standing stock. 相似文献
The Agulhas Current with its retroflection and attendant eddy-shedding is the cause of some of the greatest mesoscale variability in the ocean. This paper considers the area to the south and east of Madagascar, which provides some of the source waters of the Agulhas Current, and examines the propagating sea surface height signals in altimetry and output from a numerical model, OCCAM. Both show bands of variability along the axis of the East Madagascar Current (EMC) and along a zonal band near 25°S. Sequences of images plus associated temperature data suggest that a number of westward-propagating eddies are present in this zonal band. The paper then focuses on the region to the south of the island, where ocean colour and infra-red imagery are evocative of an East Madagascar Retroflection. The synthesis of data analysed in this paper, however, shows that remotely observed features in this area can be explained by anticyclonic eddies moving westward through the region, and this explanation is consistent with numerical model output and the trajectories of drifting buoys. 相似文献
A three-level procedure for assessing jack-up foundation stability for more or less homogenous soils is described. The objective is to provide a rational framework for these assessments that ensures their safe operation in extended year-round operations and enables their use in deeper waters than at present.
The three levels of the procedure have to be entered successively as long as foundation stability cannot be proven. The first level is a screening exercise and essentially replaces the well-known preload check. The second level compares factored foundation loads resulting from a structural analysis with foundation capacities obtained with ultimate bearing capacity formulae. The most refined third level assesses whether the displacements associated with these loads lead to an acceptable situation, i.e. capacity increase and/or load redistribution that does not result in collapse of the jack-up unit.
Since, for maximum benefit, this third-stage analysis requires a non-linear foundation model to be linked with the structural package used: such a tool is provided in the paper. Examples are given to demonstrate the impact of the assessment procedure.
This procedure forms part of the overall in-house approach to the assessment of jack-ups and has already been offered to the jack-up industry as part of the continuing efforts towards establishing common and accepted standards for jack-up assessments. Further developments have been identified and will be pursued. 相似文献
Sandy sediments in shallow coastal waters of the Baltic Sea are often characterised by large numbers of biogenic structures which are produced by macrozoobenthos species. A series of experiments was devised to quantify how the interaction of such structures with the near-bed flow regime affects the sediment flux. Most experiments were done with simplified replicates of structures generated by typical species commonly found in the Mecklenburg Bight, starting with solitary structures and regularly-spaced arrays in a range of characteristic population densities, followed by a complex benthic macrofauna community, both artificial and alive. A laboratory flume channel, equipped with an acoustic Doppler flow sensor and a topography scanning laser, was used for high-resolution measurements (2 mm horizontal step size and 0.3 mm vertical resolution) of sand erosion (220 µm median grain size, at 20 cm s− 1) and fine particle deposition (8 µm grain size, at 5 cm s− 1). Sediment transport threshold values were measured for each layout. As a rule-of-thumb, both the erosion fluxes and the deposition of suspended matter increased considerably at low population densities (below 2%, expressed as percent of the sediment surface covered, i.e. roughness density RD). Above densities of 4%, erosion almost stopped inside the test arrays, and deposition remained well below the level of unpopulated areas. An attempt to extrapolate these findings to field conditions (using field current velocity data from 2001) showed that the net flux switched from erosion to deposition for densities above 5%. These parameters can now be integrated into a numerical sediment transport model coupling waves, currents, sediment dynamics and biological processes, which is currently under construction at the Baltic Sea Research Institute (IOW), Rostock, Germany. 相似文献
During the period 1992–1998, 38 isobaric RAFOS floats were deployed to sample the subsurface flow of the California Undercurrent. The deployments, released over the California continental slope west of San Francisco, have sampled robust year-round poleward subsurface flow associated with the Undercurrent most seasons and the combined inshore current and Undercurrent in winter. Two other types of flow have been seen: a region of weak flow with little net displacement just west of the California Undercurrent, and an active westward propagating eddy field. This eddy field appears to be the primary mechanism for moving floats from the Undercurrent into the ocean interior. The observations and statistics from the RAFOS floats are compared with Lagrangian estimates of particles tracked in a global high resolution ocean simulation in order to evaluate the fidelity of the model along an eastern boundary. The results show that the model reproduces the general character of the flow reasonably well, but underestimates both the mean and eddy energies by a substantial amount. 相似文献
Estuaries are last in line as a recipient of river water and for this reason they are particularly vulnerable to negative environmental impacts due to water scarcity and pollution. They only receive the runoff that has not been abstracted or prevented from reaching rivers. When this runoff is substantially reduced their functionality is undermined and they often become less attractive for recreational use. This article explores some aspects entailed in efficiently managing the allocation of water to estuaries problem and some associated problems. It is shown that efficient management requires the marginal social costs of the inflows to be brought into equivalence with the marginal social values of the inflows, and these values may be estimated, but that there are the challenges in this estimation and in linking these estimates to the welfare of the people in whom the managers of river systems are (presumed to be) interested. 相似文献
Abstract Increasing concentrations of carbon dioxide and other gases are expected to warm the earth several degrees in the next century by a mechanism known as the greenhouse effect. Such a warming could cause sea level to rise two to five feet by expanding ocean water, melting mountain glaciers, and perhaps eventually causing polar glaciers to melt and slide into the oceans. A rise in sea level of even three feet could cause substantial erosion of beaches and coastal wetlands, increased flooding, and intrusion of saltwater into rivers, bays, and aquifers. Fortunately, many of the adverse consequences can be avoided by taking timely measures in anticipation of sea level rise. Nevertheless, many coastal zone managers are reluctant to take these measures until the prospect of sea level rise becomes more certain. This article examines the implications of future sea level rise and identifies anticipatory measures that may be appropriate today in spite of current uncertainties. 相似文献